Rhyticeros

Rhyticeros
Wreathed hornbill (R. undulatus)
Scientific classification
Domain:	Eukaryota
Kingdom:	Animalia
Phylum:	Chordata
Class:	Aves
Order:	Bucerotiformes
Family:	Bucerotidae
Genus:	Rhyticeros Reichenbach, 1849
Type species
Buceros plicatus[1] Latham, 1790
Species
See text.

Rhyticeros is a genus of medium to large hornbills (family Bucerotidae) found in forests from Southeast Asia to the Solomons. They are sometimes included in the genus Aceros. On the other hand, most species generally placed in Aceros are sometimes moved to Rhyticeros, leaving Aceros as a monotypic genus only containing the rufous-necked hornbill.

All species generally placed in Rhyticeros have relatively low, conspicuously wreathed casques and a mainly dull whitish horn-colored bill. Both sexes have mainly black plumage, but the head and neck of the males are white or rufous. The tail is white except in the black-tailed Sumba hornbill. They have conspicuous inflatable skin on the throat, which is blue in all except the males of the plain-pouched and wreathed hornbills, where it is yellow.

Species

The following six species are placed in the genus:^[2]

Image	Scientific name	Common Name	Distribution
	Rhyticeros plicatus	Papuan (or Blyth's) hornbill	Wallacea and Melanesia.
	Rhyticeros narcondami	Narcondam hornbill	Narcondam
	Rhyticeros subruficollis	Plain-pouched hornbill	southern Myanmar, adjacent parts of western Thailand and northern Peninsular Malaysia
	Rhyticeros undulatus	Wreathed hornbill	north-eastern India and Bhutan, east and south through mainland Southeast Asia and the Greater Sundas in Indonesia
	Rhyticeros everetti	Sumba hornbill	Sumba in the Lesser Sunda Islands
	Rhyticeros cassidix	Knobbed hornbill	Sulawesi, Buton, Lembeh, Togian and Muna Island.

An undescribed extinct hornbill species from Lifou in the Loyalty Islands, living until at least some 30,000 years ago, was initially placed in Aceros, but its biogeography places it with the species now in Rhyticeros (Steadman, 2006).

References

• Kemp, A. C. (2001). Family Bucerotidae (Hornbills). pp. 436–523 in: del Hoyo, J., Elliott, A., & Sargatal, J. eds. (2001). Handbook of the Birds of the World. Vol. 6. Mousebirds to Hornbills. Lynx Edicions, Barcelona. ISBN 84-87334-30-X
• Steadman, David William (2006): Extinction and Biogeography of Tropical Pacific Birds. University of Chicago Press. ISBN 0-226-77142-3

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Genera of trogons, hornbills, kingfishers, woodpeckers and their extinct allies

• Kingdom: Animalia
• Phylum: Chordata
• Class: Aves
• Infraclass: Neognathae
• Clade: Coraciimorphae
• Clade: Cavitaves

Cavitaves
Leptosomiformes Leptosomidae Leptosomus †Plesiocathartes? †Waltonavis Eucavitaves Trogoniformes Trogonidae Apalharpactes Apaloderma †Eotrogon Euptilotis †Foshanornis Harpactes †Masillatrogon †Paratrogon Pharomachrus †Primotrogon Priotelus †Septentrogon Trogon Picocoraciae Bucerotiformes See below ↓ Picodynastornithes Coraciiformes See below ↓ Piciformes See below ↓

Bucerotiformes
Bucerotiformes †Laurillardiidae Laurillardia †Messelirrisoridae Messelirrisor Upupi Phoeniculidae †Phirriculus Phoeniculus Rhinopomastus Upupidae Upupa Buceroti Bucorvidae Bucorvus †Euroceros Bucerotidae Aceros Anorrhinus Anthracoceros Berenicornis Buceros Bycanistes Ceratogymna Horizocerus Lophoceros Ocyceros Penelopides Rhabdotorrhinus Rhinoplax Rhyticeros Tockus

Coraciiformes
incertae sedis †Cryptornis †Paracoracias †Quasisyndactylus †Eocoraciidae Eocoracias †Geranopteridae Geranopterus †Primobucconidae Primobucco Septencoracias Brachypteraciidae Atelornis Brachypteracias Geobiastes Uratelornis Coraciidae †Chehuenia Coracias Eurystomus †Miocoracias Meropidae Meropogon Merops Nyctyornis Alcedines Alcedinidae Alcedininae Alcedo Ceyx Corythornis Ispidina Cerylinae Ceryle Chloroceryle Megaceryle Halcyoninae Actenoides Caridonax Cittura Clytoceyx Dacelo Halcyon Lacedo Melidora Pelargopsis Syma Tanysiptera Todiramphus Momotidae Aspatha Baryphthengus Electron Eumomota Hylomanes Momotus †Protornis? Todidae †Palaeotodus Todus

Piciformes
incertae sedis †"Homalopus" †Gracilitarsidae Gracilitarsus Neanis †Sylphornithidae Oligosylphe Palaegithalus? Sylphornis Galbuli †Jacamatia Bucconidae Bucco Chelidoptera Hapaloptila Hypnelus Malacoptila Micromonacha Monasa Nonnula Notharchus Nystalus Galbulidae Brachygalba Galbalcyrhynchus Galbula Jacamaralcyon Jacamerops Pici incertae sedis †Rupelramphastoides †Miopiconidae Miopico †Picavidae Picavus Ramphastides Capitonidae Capito †Capitonides Eubucco Lybiidae Buccanodon Cryptolybia Gymnobucco Lybius Pogoniulus Pogonornis Stactolaema Trachyphonus Tricholaema Megalaimidae Caloramphus Psilopogon Ramphastidae Andigena Aulacorhynchus Pteroglossus Ramphastos Selenidera Semnornithidae Semnornis Picides Indicatoridae Indicator Melichneutes Melignomon Prodotiscus Picidae See below ↓

Picidae
incertae sedis †Bitumenpicus †Breacopus †Palaeonerpes †Palaeopicus †Piculoides Jynginae Jynx Picumninae Picumnus Sasiinae Sasia Verreauxia Picinae †Australopicus †Palaeonerpes †Pliopicus Nesoctitini Nesoctites Hemicircini Hemicircus Picini Campethera Celeus Chrysophlegma Colaptes Dinopium Dryocopus Gecinulus Geocolaptes Micropternus Meiglyptes Mulleripicus Pardipicus Piculus Picus Campephilini Blythipicus Campephilus Chrysocolaptes Reinwardtipicus Melanerpini Chloropicus Dendrocopos Dendrocoptes Dendropicos Dryobates Leiopicus Leuconotopicus Melanerpes Picoides Sphyrapicus Veniliornis Xiphidiopicus Yungipicus